Penn State redshirt freshman quarterback Ethan Grunkemeyer threw for 219 yards and a touchdown in the Lions' 27-24 loss to Indiana on Saturday.

The Hoosiers score the game-winning touchdown with 36 remaining, sending Penn State to its sixth straight loss.

The Hoosiers' hurry-up in front of 100,000-plus at Beaver Stadium culminated with Mendoza's 7-yard toss to Omar Cooper Jr. in the back of the end zone, who dazzled despite solid coverage and came down with the football with his left toe inbounds to push Indiana ahead.

After a Penn State punt and a 7-yard sack, Indiana moved downfield behind Fernando Mendoza, who completed five passes for 87 yards. He threw a high pass to the back of the end zone for Cooper, who leaped and caught it over Zakee Wheatley.

But then it collapsed. Indiana quarterback Fernando Mendoza made one spectacular play after another and the Hoosiers ripped Penn State’s heart out, winning the game on a beautiful throw by Mendoza to Omar Cooper Jr. The air was sucked out of Beaver Stadium.

Penn State had No. 2 Indiana on the ropes at Beaver Stadium as time ticked down, but an astounding toe-tap touchdown from Omar Cooper Jr. kept the Hoosiers' undefeated season alive.

STATE COLLEGE | After video replay confirmed Indiana receiver Omar Cooper Jr.’s touchdown catch, Penn State interim head coach Terry Smith was fuming and leaned into the officiating crew. It was the most anger and frustration Smith has shown on the sideline in his four weeks on the job.

Ethan Grunkemeyer and the Lions' passing attack improved greatly, but an overall strong defensive performance came up just short.